On Cybertron, Vector Sigma has become stable again after the fierce fighting between the Cybertrons and Destrons came to an end. The planet became peaceful again. Vector Sigma has created something that could help the Cybertrons greatly.
However, Soundblaster and his cassettes have managed to get onto the planet, and have discovered what Vector Sigma is creating. Soundblaster withdraws from Cybertron and heads back to Jaar, where he informs Galvatron on what is going on. Vector Sigma is making something called "Cybertonuron", which is an alloy many times stronger than Cybertonium. Galvatron thinks it sounds amazing, and wants to get a hold of it.
The reason for this is because he could then create a Cybertonuron sword, and use it to kill Rodimus Convoy and destroy Fortress' ship. Galvatron likes these ideas a lot, as this Cybertonuron would make the Destrons invincible.
Soundblaster goes on to tell Galvatron about how this new metal can only be created by Vector Sigma itself and that if the Destrons got a hold of it they could conquer the entire universe. Galvatron decides the only way they can get it is to invade Cybertron itself. Meanwhile, Twincast has found out about Cybertonuron and reports it to Rodimus. Elsewhere, the Cybertron Headmasters are working out, but Chromedome is becoming anxious to fight and to take the battle to the Destrons. The others don't agree and say being hot-headed can lead to trouble. Chromedome regrets opening his mouth.
Back on Jaar, Galvatron has the Animatrons sent to Cybertron to start things off. As Galvatron walks outside, Scorpnok appears. Scorponok isn't with the plan and thinks Galvatron is taking a risk, as there is no guarantee that this new alloy can even be completed yet. He doesn't want their side to make sacrifices in vain. Galvatron tells him that he'll decide what is in vain and tells Scorponok to get ready.
Galvatron, Soundblaster, Scorponok and the Destron Headmasters head to the Earth Destron base via Astrotrain. As they land, Galvatron and Soundblaster run straight inside the base. The Destron Headmasters are about to follow when Scorponok tells them to wait. He says he is against Galvatron's plan, and they need to be mindful of what he is letting them in for.
As the Destron Headmasters head inside the base, their movements have been seen by some of Twincast's cassettes. They return to Twincast, who reports to the Cybertrons on Athenia. He doesn't know what the Destrons are up to, but it doesn't look good. Chromedome says he'll go to Earth to find out more, and Hardhead wants to go with him, but Fortress tells them they must go only to do some reconnaissance -no fighting yet.
Meanwhile the Animatrons have arrived on Cybertron, and as they check their weapons, a couple of Cybertron troops spot them. One has the not so bright idea of walking straight towards them, and gets blasted. He tells the other to go and contact Rodimus Convoy quickly. Meanwhile, Galvatron has rounded up the Stuntrons and Combatrons, and prepares to head to Cybertron.
Meanwhile, at the Destron base, Sixshot reports to Scorponok something sinister -Galvatron has told Sixshot that he intends to banish Scorponok after he gets the Cybertonuron. He suggests Scorponok just do as Galvatron wants, but Scorpnok refuses, saying that he has always worked for the good of the Destrons. Sixshot says the alloy has not been completed yet, but if they could get it they could crush their enemies. Outside the base, Chromedome and Hardhead arrive, but are surprised to see nobody around, not even any guards. This is become most of the Destrons have already arrived on Cybertron via the space bridge.
Galvatron tells everyone to move forward, and soon he and the Destrons with him are causing massive damage to the planet. Galvatron says Cybertron will soon belong to him.
Some Cybertron troops fire back, but are no match for the Destrons and most of them get killed pretty quickly. One manages to run off and contact Rodimus Convoy on Athenia.
When Rodimus learns that Cybertron is under attack, he tells Arcee to contact Ultra Magnus on Earth and have him send reinforcements to Cybertron via the space bridge. Rodimus is going to go to Cybertron also, but as he prepares to leave, Fortress talks to him. Fortress wants him to hold back their attack, as Galvatron expects the Cybertrons to show up. He thinks they should come up with another plan, but Rodimus does not agree. Fortress says he knows this new alloy is important, but they still don't know if it can actually be created yet. Rodimus tells him that it isn't just the Cybertonuron -Cybertron is their homeworld, they cannot just hand it over to the Destrons. He still intends to defend it.
On Cybertron City on Earth, the Trainbots, Airbots and Protectbots are about to depart to Cybertron. Chromedome and Hardhead arrive and decide to go with them, but check in with Fortress first. Brainstorm is the one who answers, and Chromedome just tells him he and Hardhead are going to Cybertron, they'll see them there. Fortress tells him not to be rash, but Chromedome ends the communication. Fortress decides they'd better get moving. Rodimus, Arcee, Blurr and Chear get onboard the ship and it blasts off seconds later.
Meanwhile, on Cybertron, the battle is largely one sided as the Cybertron guards are just useless against the Destrons. Galvatron thinks it is natural for them to be losing, but then the Earth reinforcements arrive. Chromedome and Hardhead fight their Destron counterparts...
...whilst the various special teams combine and fight each other. Fortress' ship reaches Cybertron, and Rodimus jumps out of it. He finds Galvatron and says this is the last time they will fight, as he intends to end it all here. Galvatron laughs and thinks he is talking nonsense. This planet will be his. The two start to battle.
Brainstorm and Highbrow see that Chromedome and Hardhead are in trouble, so they also jump out of their ship and help them. Meanwhile, Galvatron punches Rodimus so hard he crashes into a wall. Galvatron then transforms and starts shooting at the building behind Rodimus.
Part of the wall falls away and the debris covers Rodimus. Galvatron says he'll finish him off, but then Soundblaster comes over. He asks Galvatron to step aside with him. Galvatron tells him to just tell him whatever it is he has here, but Soundblaster says he does not want the Cybertrons to hear this information. Galvatron reluctantly follows him, giving Blurr, Chear and Arcee the chance to help Rodimus.
Soundblaster's important news is that Vector Sigma has finished research on Cybertonuron and that production could start at any moment. Galvatron says this is perfect, all they need to do is crush the remaining Cybertrons. Meanwhile, Scorponok has also found out about the Cybertonurons completion, and believes they cannot afford the slightest mistake now. On his ship, Fortress is still thinking about what he told Rodimus earlier, and wants to end this battle.
Scorponok thinks about the worst-case scenario -if the Destrons lost and the Cybertrons got the new alloy. He thinks that Fortress could put the alloy all over his ship, and say if Galvatron fired at it, his shot would just bounce off and end up hurting their own forces.
Scorponok thinks this is not good and knows they cannot let the Cybertrons get the Cybertonuron. There is only one option to guarantee this now -the destruction of Vector Sigma and the destruction of planet Cybertron itself. He calls out to Weirdwolf, Skullcruncher and Mindwipe.
The three Destrons have some kind of telepathic link to him, and hear his words. They knock the Cybertron Headmasters back and tell them they will continue their battle later. They find Scorponok, who tells them how they will crush them all in one go. He reveals three powerful bombs, which they will use to blow up Cybertron.
Mindwipe is to set the bombs up, whilst Skullcruncher and Weirdwolf are to support him. They will have enough time to get away before the bombs explode, and he will make sure the other Destrons are off the planet. Will the Destrons succeed? And what will happen to the Cybertrons?
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